Gungans (Second Human Empire name Gujj'levvel Ryiink'al) are a semi-aquatic humanoid species indigenous to planet Naboo. Gungans are tall humanoids with a bill-like mouth, large nostrils which close when they are diving, and very long ears which aid in swimming. Their eyes have nictitating membranes (transparent eyelids) in order to see underwater. The Otolla subspecies is leaner, have a long neck and short partially-retractable eyestalks, unlike the stout Ankura subspecies. Gungans are omnivorous and have strong teeth and a long extendable tongue used to capture food. Some Otolla Gungans develop short catfish whiskers as they age.